Geology museums of mainland Europe: The Museo di Storia Natural, geology and palaeontology, Verona

Jon Trevelyan (UK)

The geology and palaeontology section at the Museo di Storia Naturale: geology and palaeontology in Verona is one of those museum experiences that perhaps quietly insists you pay attention if you notice it there among the other attractions of this lovely town.

Fig. 1. The entrance to the museum.

It doesn’t shout with flashy interactives or staged drama (which I hate). Rather; it offers volume and provenance – rooms of specimens whose value is more in the sum of things than in a single blockbuster display. That solidity is both the exhibit’s great strength and, frustratingly, its central curatorial problem: an embarrassment of riches that is not always given the contemporary presentation it needs.

Fig. 2. A cabinet of Bolca fish, including the barracuda in Fig. 39.
Fig. 3. Zanthoxylon is a genus of spiny, aromatic trees and shrubs in the citrus family (Rutaceae), found today in tropical and subtropical regions worldwide. They have pinnate leaves, prickly stems, small clustered flowers and peppery seeds. Fossil species are known from Cenozoic deposits and indicate warm, tropical climates in the past.

Housed in the historically resonant Palazzo Pompei with deposits also at the Arsenal, the geology and palaeontology section is part of a civic collection built over centuries. The museum’s own figures emphasise scale. The holdings for the section are estimated in the hundreds of thousands, with a seriousness of collection that places Verona comfortably among those regional natural-history repositories with scientific force. You feel that history the moment you enter the imposing atrium (Fig. 4), and then with the wooden casework, dense glass cabinets, drawers, and the slow, reassuring thud of academic accumulation rather than tourist theatre.

Fig. 4. The atrium to the museum. It provides an awe-inspiring entrance.

If there is a single headline to the collection, it is the Bolca fossils and, boy, are they good! Bolca is a rich Eocene lagerstätte from the Val d’Alpone area, and is world-famous for exquisitely preserved fishes and plants, and Verona’s palaeontology rooms understandably make this local heritage the centrepiece.

Fig. 5. Eoscatophagus frontalis is an extinct, Eocene bony fish from Monte Bolca that is laterally compressed and round-bodied, it resembled modern scat fishes and lived in warm, shallow marine environments.

The museum reportedly conserves thousands of Bolca fish specimens (the broader collections apparently include some 2,500 identified Bolca fish in older inventories), alongside plant fossils that hint at a much warmer, near-tropical past.

Fig. 6. Beautiful palms from the Bolca region.
Fig. 7. More plants from the Bolca region.

These displays are not only beautiful but scientifically significant – they tell a story about Eocene marine ecosystems and the Alps’ deep history. I was astonished by them.

Fig. 8. Various gastropods and bivalves.

Beyond Bolca, there are regional and Pleistocene collections worth noting. There are cave deposits, Quaternary vertebrate remains, and a wide array of invertebrate and plant fossils from local sites such as Monte Vegroni, Monte Postale and Negrar. For visitors with an interest in palaeobiology, the museum’s range – vertebrates, invertebrates and plants – is impressive and gives a useful, if somewhat old-school, panorama of the area’s fossil record.

Fig. 9. A video of the Bolca exhibition room.

But while the museum’s collections are rich, the exhibition design feels rooted in the nineteenth or mid-twentieth century. I don’t mind this. Many cabinets are filled densely, labels are short (and mostly in Italian), and the lighting – often low and uneven – does little to accentuate the specimens’ aesthetic or scientific value.

Fig. 10. Another palm from the Bolca region.

But I’m willing to forgive this, as I love such old-fashioned museums. So for me, it’s a minor quibble that, on the whole, fossils, minerals and thin-section displays all benefit from thoughtful lighting and clear labelling. In several rooms, you can tell the curators know the best specimens, but the visitor cannot always see this, as they are buried in crowded glass cabinets and small print.

Fig. 11. Sparnodus vulgaris is an extinct species of fish from the Eocene found in the Monte Bolca lagerstätte. It was a type of perciform fish belonging to the family Sparidae (sea breams) and is known from fossils discovered in the Monte Bolca Limestone.

However, strip away these curatorial gripes and you still have a working scientific archive. The museum’s holdings are used in research; and the collections include type material and regionally important specimens.

Fig. 12. Caesalpinia eocenica is an extinct species of leguminous plant (in the Fabaceae family, subfamily Caesalpinioideae) that lived during the Eocene epoch.

The Bolca assemblage alone is something field workers and ichthyologists will still travel to study. The geographic proximity of the specimens to the sites where they were discovered also gives the museum educational leverage: local geology students and amateur fossil collectors can see the direct connection between field sites and museum curation. In short, the section remains a true research resource, not simply an old-fashioned cabinet of curiosities.

Fig. 13. Guajacites sp is an extinct Eocene fossil wood or fruit from Monte Bolca, resembling modern tropical hardwoods (Guaiacum). It indicates the presence of warm, subtropical coastal forests.

So, bring your patience and your curiosity. The museum rewards the careful, the inquisitive, and those who enjoy the quiet pleasure of a dense, well-provenanced cabinet. And if you go specifically for Bolca, don’t be surprised if you leave thinking about seas that existed where hills now stand. The fossils are mainly from the wider Verona region and nearby Veneto fossil sites, and there’s a clear hierarchy in what’s scientifically or visually most striking. A don’t miss out on the mineral collections and more recent animal and human collections.

Fig. 14. It’s not just fossils. Here are some cabinets of minerals.

The Bolca fish fossils (Eocene)

Bolca, in the Lessini Mountains about 30km east of Verona, is one of the world’s most famous fossil sites – a Lagerstätte, but note that there is little in the way of opportunities to actually visit these sites. However, the Verona museum holds a large portion of its best material.

Fig. 15. Ophidium voltianum is an extinct Eocene cusk-eel-like fish from Monte Bolca. It has; elongated and slender-bodied and lived near the sea floor in warm, shallow marine habitats.

There are perfectly articulated fish skeletons on fine limestone slabs, which look almost painted, but the detail is real – spines, scales and even soft-tissue impressions. There is also a wonderful variety of species, for example, Priscacara, Eoholocentrum, Ductor, Caranx and Sphyraena (an ancient barracuda; Fig. 39). Some resemble modern reef fish, showing that Bolca once lay in warm, tropical seas. Fossil plants (palm fronds, leaves) sometimes occur on the same bedding planes as the fish, suggesting a near-shore, lagoonal environment.

Fig. 16. Orrichthys longimanu (left and right), Histionotphorus bassanii (centre) and Eophrnie barbutii (bottom right). O. longimanus is an extinct Eocene batfish from Monte Bolca, Italy; deep, laterally compressed body with long fins, adapted to warm, shallow reef environments. H. bassanii is an extinct Eocene sailfish-like fish from Monte Bolca. It has an elongated body and a prominent dorsal fin, making it a fast-swimming predator in shallow tropical seas. E. barbutil is an extinct Eocene reef fish from Monte Bolca, laterally compressed with long fins, which lived in shallow tropical seas feeding on small invertebrates.

And the contrast between the beige limestone and the dark fossils is stunning. Note how some are displayed under raking light so you can see relief. The Bolca room is the place to linger and, if you see anything labelled ‘Pesci di Bolca’ or ‘Monte Bolca,’ stop – these are specimens of global significance.

Fig. 17. Psettopsis subarcatus is an extinct species of bony fish from the Eocene deposits of Monte Bolca. It belonged to the flatfish-like order Perciformes, with a laterally compressed body adapted for life near the sea floor.

Fossil plants from Monte Bolca and Monte Postale (Eocene)

Plant fossils, from the same strata as the fish, record the vegetation of an early tropical lagoon environment (largely from the Monte Vegroni, Palmeto Cerato quarries). And it is these that perhaps I loved the most.

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Fig. 18. More superb palms from the Bolca region.
Fig. 19. Latanites praticinensis is an extinct Eocene fan palm from Monte Bolca and Praticino, with large palmate leaves indicating that it grew in warm, coastal, subtropical forests.

There are palm leaves and mangrove-like plants (look for elongated, segmented fronds). And there are charred fragments or compression fossils that are surprisingly dark, which tell you about oxygen-poor burial conditions that allowed preservation.

Fig. 20. Depanocarpus docampii is a fossil fruit or seed species from the Eocene deposits of Monte Bolcay. It represents a tropical or subtropical tree fruit, likely related to modern Malvaceae or Sterculiaceae-type plants.
Fig. 21. Dombeyopsis hibiscifolia is an extinct Eocene tropical tree from Monte Bolca; known from broad, heart-shaped leaves, it inhabited warm coastal forests and contributes to reconstructing subtropical Eocene vegetation.

Some specimens are labelled with ‘Lauraceae’ or ‘Arecaceae’ (laurel and palm families). These plants suggest the Verona region was once as warm as modern Indonesia.

Fig. 22. Castellinia compressa is a fossil species of ‘pandanei-like’ or palm-fruit type remains from the Eocene of northern Italy.
Fig. 23. Fracastoria melo is a fossil fruit species from the Eocene deposits of Monte Bolca, described by Massalongo in the nineteenth century. It represents a large, rounded fruit resembling those of modern tropical trees, possibly related to the Sterculiaceae or Malvaceae families, and indicates a warm, subtropical climate in Eocene northern Italy.

Marine invertebrates – ammonites, nautiluses and belemnites (Jurassic and Cretaceous)

Before the Alps rose, this part of Italy lay under shallow seas teeming with shelled cephalopods. So, there are copious ammonites, sometimes polished to show internal chambers. They’re from local limestone formations, possibly the Rosso Ammonitico Veronese – a famous red limestone used in architecture (Fig. 24).

Fig. 24. Phylloceras sp. These are abundant in the Oxfordian–Tithonian part of the Rosso Ammonitico sequence. The ammonite was embedded in the wall of a restaurant near the monastery of Santuario Madonna della Corona.

Other fossil cephalopods include belemnites – the bullet-shaped internal skeletons of extinct squid relatives.

Note the differences in preservation – some are calcitic casts, others are filled with crystalline spar. And if you notice an ammonite labelled ‘Rosso Ammonitico,’ look closely at the rock – that red matrix is the same stone you see in many Verona buildings and pavements (Figs. 25 to 27 ).

Fig. 25. A large ammonite on the pavement of Verona with drain for scale.
Fig. 26. Ammonites on the pavements of Verona.
Fig. 27. An ammonite in the pavement with my feet for scale (size 9).
Fig. 28. Nautilus disculus is an extinct species of nautiloid cephalopod from the Late Jurassic period.

Note also the bivalves, including pectens (Fig. 29) from the Carboniferous and scallops (Fig. 30) from the Miocene to Quaternary.

Fig. 29. Pecten incrassatus is an extinct species of fossil mollusc, specifically a type of scallop or bivalve, known from the Carboniferous period.
Fig. 30. Pecten latissimus is an extinct species of fossil scallop shell from the Miocene to Quaternary.

Quaternary (Ice Age) mammals from local caves and river terraces (Pleistocene)

These fossils connect directly with Ice Age fauna of northern Italy – cave bears, mammoths and rhinos. There are cave bear (Ursus spelaeus) skulls and limb bones, often from the Lessini caves (Fig. 31), together with mammoth (Fig. 32) and woolly rhinoceros teeth – enormous molars with ridged chewing surfaces.

Fig. 31. Cave bear (Ursus arctos) from the Late Pleistocene L’Orso delle Caverne.
Fig. 32. Woolly mammoth remains and the Mammoth in Fig. 36.

And there are deer antlers and horse bones, sometimes with stratigraphic labels showing their provenance from river gravels. The labels might say ‘Quaternario’ or ‘Pleistocene’. Many were collected in the nineteenth century when speleology in Verona was fashionable.

Fig. 33. Some slightly more modern exhibits at the museum.

Fossil corals and reef limestones (mainly Jurassic to Eocene)

The fossils at the museum illustrate how the region’s ancient seas built reef structures long before the modern Mediterranean. There are colonial coral heads – rounded masses with hexagonal surface patterns, and single ‘horn corals’ (Rugosa) in older beds. You can often see that the rocks show growth laminae or the internal septal pattern – these were living reefs. These fossils link to the limestones that later became building stones of Verona, with geology meeting architecture again.

There is also an extinct Late Cretaceous sea turtle from northern Italy, likely related to the leatherback lineage (Fig. 34).

Fig. 34. Protosphargis veronensis is an extinct Late Cretaceous sea turtle from northern Italy, known from carapace fragments (but this specimen is superb). It was a small-to-medium sized pelagic turtle living in shallow Tethys Sea waters.

Vertebrate bones from the Verona area (Miocene to Pleistocene)

Scattered through the section are fossils of terrestrial vertebrates that track the transition from the Eocene (Fig. 35) warm Miocene ecosystems to colder Pleistocene ones (Fig. 36).

Fig. 35. Asiatosuchus depressifrons is an extinct species of crocodile that lived in Europe during the early Eocene.
Fig. 36. Mammuth cf. americanus (American mastodon). Extinct Pleistocene elephant from North America, which was large-bodied, with long, curved tusks and a thick coat, adapted to cold steppe and woodland environments.

There are also fossilised bones and jaws of small mammals, sometimes fragmentary, including Hipparion (a three-toed horse).

Microfossils and thin sections

Though easy to overlook, the glass slides and microfossil displays show how palaeontologists identify strata and date rocks. There are small vials or microscope slides labelled ‘foraminiferi’ (tiny single-celled marine organisms), and thin sections showing cross-sections of corals, shells and microfauna. Sometimes, a nearby microscope lets you view them magnified.

Geological context and the link to Verona stone

A fascinating aspect of the museum is the cases with three dimensional palaeontological maps (Fig. 37) showing distribution of finds across Verona and the regional stratigraphy showing where the relevant formations occur.

Fig. 37. One of the remarkable three=dimensional geological maps of the area.

These give you a sense of geological context and are part of the museum’s efforts to tie fossils to geology, showing that Verona’s famous building stones are literally made of fossil-rich limestone. In this context, there are display cases of ‘Rosso Ammonitico Veronese’, ‘Nembro Limestone’ and ‘Scaglia Rossa’, with embedded fossil shells, ammonites and crinoids in the slabs.

Fig. 38. Sphyaena prisca (barracuda) is an extinct Eocene barracuda-like fish from Monte Bolca, Italy; elongated and streamlined, and was a fast-swimming predator in warm, shallow marine waters.

Historical collections and early palaeontological labels

The museum preserves specimens collected since the 1700s. There are old handwritten labels with ornate script, often giving nineteenth-century localities and early display drawers showing how fossils were classified before Darwin. Sometimes, you’ll find type specimens or references to collectors like Santi or the Bolla brothers, which together display what could be called the ‘evolution of palaeontology itself’, that is how fossils moved from ‘curiosities’ to data.

Fig. 39. Sphyaena prisca (barracuda) is an extinct Eocene barracuda-like fish from Monte Bolca, Italy; elongated and streamlined, and was a fast-swimming predator in warm, shallow marine waters.
Fig. 40. Eorandallius elegans (bottom left) and Pesce chirugo giallo (top). E. elegans is an extinct Eocene boxfish from Monte Bolca. Deep-bodied with a bony, box-like carapace, it inhabited warm, shallow reef environments. And P. chirurgo giallo is an extinct Eocene yellow surgeonfish from Monte Bolca. Laterally compressed with sharp tail spines, it lived in warm, coral-rich coastal waters.

Final thoughts

The Verona Natural History Museum’s fossil galleries aren’t ‘wow’ in a modern theatrical sense. But if you go in prepared, you’ll see one of Europe’s most important local fossil collections. It’s the kind of place where understanding multiplies appreciation. That is, knowing what Bolca represents or how Rosso Ammonitico underpins the city’s architecture, turns an unlabelled slab into a time capsule.

Fig. 41. Eoplatax subvespertilio is an extinct Eocene spadefish-like bony fish from Monte Bolca, Italy; laterally compressed and deep-bodied, it inhabited shallow, reef-associated marine environments.

Go slowly, question what you see, and imagine Verona shifting from coral reef to lagoon to Ice Age steppe. That’s what the fossils are really showing – and that’s what most visitors miss because the labels don’t spell it out.

Fig. 42. Galeorhinus cuvieri is a shark found mainly in Eocene and Oligocene marine deposits of Europe, especially the Monte Bolca site.
Fig. 43. Ceratoichtys pinnatiformis is an extinct Eocene bony fish from Monte Bolca.
Characterised by a deep. Also with a laterally compressed body this time with pronounced fins, it lived in warm, shallow reef environments.
Location
The museum is located in Palazzo Pompei on the Lungadige (an atmospheric stretch along the Adige) and the Natural History Museum’s opening hours are published on the museum’s website (see ab0ve). Typically open Tuesday through Sunday, with the usual museum rhythm of a last entrance before closing.
Visiting the museum
If you’re visiting Verona and care about Bolca or regional geology, plan to give the museum at least an hour, preferably two – one to browse and get the lay of the land, another to go back and focus on Bolca and the palaeontological cabinets. Expect many labels in Italian and rooms that favour content over spectacle.
Other articles in this series
Geology museums of mainland Europe: A hidden gem in the heart of Barcelona – the Geological Museum of the Seminary, by Paul D Taylor (UK) and Consuelo Sendino (UK)
Geology museums of mainland Europe: Museo di Geologia e Paleontologia, Florence, by Jon Trevelyan (UK)
The Geology museums of mainland Eurpe: The Siatista Historical Paleontological Collection, Greece – the first record of a stegodon in Europe and the making of the straight-tusked elephant, by Dick Mol (The Netherlands), Evangelia Tsoukala (Greece), Evangelos Vlachos (Greece), Anna Batsi (Greece), Hans Wildschut (The Netherlands), Dimitra Labretsa (Greece) and Wilrie van Logchem (The Netherlands)
Geology museums of mainland Europe: Highlights from the Museum am Löwentor in Germany, by Jack Wilkin (UK)
Geology museums of mainland Europe: Musée-Parc des Dinosaures (Dinosaur Museum-Park), Mèze, France, by Fred Clouter (UK)
Geology museums of mainland Europe: The palaeontological collection at the Teyler’s Museum, Haarlem, The Netherlands, by Mike Howgate FLS (UK)
Geology museums of mainland Europe: Agios Georgios Cave, Kilkis: 50 years of history, 30.000 years of prehistory, by Vassilis Makridis (Greece), Evangelia Tsoukala (Greece), Evangelos Vlachos (Greece), Katerina Tsekoura (Greece), Wilrie van Logchem (The Netherlands) and Dick Mol (The Netherlands)
Geology museums of mainland Europe: The Museum national d’Histoire naturelle’s Galerie de Géologie et de Minéralogie, Paris, by Jon Trevelyan (UK)
Geology museums of mainland Europe: The Galerie de Paléontologie et d’Anatomie Comparée, Paris, by Jon Trevelyan (UK)
Geology museums of mainland Europe: The Naturmuseum Südtirol Bolzano, Austria, by Jon Trevelyan (UK)
Geology museums of mainland Europe: The Museo di Storia Natural, geology and palaeontology, Verona, by Jon Trevelyan (UK)

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